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Sperm route to fertilization

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To do so, the sperm cell must pass through a long and challenging path. Fig 1. The sperm cells gain their full ability to swim when they are ejaculated into the reproductive tract [1] , [2]. Upon ejaculation, the sperm cells are enclosed in a fluid called seminal plasma or semen, which is a mix of fluids from the testes, seminal vesicles, prostate, and the bulbourethral glands.
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Egg meets sperm (article) | Embryology | Khan Academy

Suarez, A. At coitus, human sperm are deposited into the anterior vagina, where, to avoid vaginal acid and immune responses, they quickly contact cervical mucus and enter the cervix. Cervical mucus filters out sperm with poor morphology and motility and as such only a minority of ejaculated sperm actually enter the cervix. In the uterus, muscular contractions may enhance passage of sperm through the uterine cavity. A few thousand sperm swim through the uterotubal junctions to reach the Fallopian tubes uterine tubes, oviducts where sperm are stored in a reservoir, or at least maintained in a fertile state, by interacting with endosalpingeal oviductal epithelium. As the time of ovulation approaches, sperm become capacitated and hyperactivated, which enables them to proceed towards the tubal ampulla.
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Molecules on the sperm's route to fertilization

The testes are where sperm is manufactured. The epididymis is a long coiled structure topping the testis, and it receives immature sperm from the testis and stores them as they mature. When ejaculation occurs, sperm are forcefully expelled from the tail of the epididymis into the vas deferens.
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In eutherian mammals billions of sperm are deposited at ejaculation in the female reproductive tract, but only a few thousand enter the oviduct. A few reach the ampulla at the time of fertilization and only one sperm fertilizes the egg. In most mammalian species the lower isthmus of the fallopian tubes has taken over the function of a reservoir in which sperm are stored under conditions that save sperm energy by suppressing motility and increase viability. Close to the time when the egg is ovulated into the ampulla, the sperm undergo a complex sequence of processes, named capacitation. Capacitation is a prerequisite for fertilization, enabling the sperm to recognize the egg and to respond to the egg signals in the appropriate manner.
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